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Cretaceous dinosaur and crocodylomorph egg records of the Sanshui Basin,Guangdong Province,southern China

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Journal of Palaeogeography 2023年第3期12卷 401-418页

作者： Li-Da Xing Zi-Sheng Guo Rui wu Chun-Lei Du Dong-Haowang w.scott persons IV State Key Laboratory of Biogeology and Environmental Geology China University of Geosciences(Beijing)Beijing100083China School of the Earth Sciences and Resources China University of Geosciences(Beijing)Beijing 100083China Grandview Museum of Natural Science Guangzhou 510630Guangdong ProvinceChina School of Earth Sciences China University of Geosciences(Wuhan)Wuhan 430074Hubei ProvinceChina Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharlestonSC 29401USA

This paper describes newly discovered dinosaur and crocodylomorph egg fragments from the Upper Cretaceous Sanshui and Dalangshan formations of the Sanshui Basin of Guangdong Province,southern *** the absence of macroscopic information,the eggshell specimens can be identified to the oofamilies Prismatoolithidae,Elongatoolithidae and Krokolithidae,and it is hypothesized that these three families of eggs correspond to troodontids,oviraptorids and crocodiloids,*** with egg fossils from Nanxiong Basin,Heyuan Basin and Ganzhou Basin highlights the similarity between these egg assemblages and those from Nanxiong *** egg fossils found in the Sanshui and the Dalangshan for-mations enrich the known faunal types in the research area and facilitate further studies on the diversity of Late Cretaceous vertebrate assemblages in the Sanshui Basin.

关键词： Prismatoolithidae Elongatoolithidae Krokolithidae Sanshui Formation Dalangshan Formation Upper Cretaceous

Largest dinosaur tracksite in China (Cretaceous, Zhaojue area, Sichuan Province): On the verge of destruction

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Geoscience Frontiers 2021年第5期12卷 124-137页

作者： Lida Xing Martin G.Lockley Hendrik Klein w.scott persons IV Guangzhao Peng Yong Ye Miaoyan wang State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing 100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing 100083China Dinosaur Trackers Research Group University of ColoradoDenver 80217USA Saurierwelt Paläontologisches Museum Alte Richt 7D-92318 NeumarktGermany Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA Zigong Dinosaur Museum ZigongSichuan ProvinceChina

The Zhaojue(ZJ)tracksites represent multiple steeply-inclined,track-bearing exposures in the clastic Feitianshan Formation of Sichuan Province that have been sequentially,excavated,mapped,expanded and destroyed,by quarrying,erosion or collapse,for almost three *** quarried area extends for more almost 1.0 km from north to south and-0.5 km from east to *** important track-bearing surfaces have been identified and identified as ZJ-Ⅰ,ZJ-Ⅱ,ZJ-ⅡN and ZJ-Ⅲ,some of which are newly exposed surfaces or expanded surfaces that were previously partially *** represent surfaces lost since they were first reported in papers published in *** ZJ-Ⅱ site represents a hugely expanded continuation of a much smaller track-bearing surface described in *** was mapped using drone technology,and now represents the largest tracksite in China,with a minimum count of 933 recognizable tracks,and the longest recorded sauropod and ornithopod trackways from China(-80 m and-52 m respectively),which also reveal a pronounced change in the direction of trackmaker *** ZJ-Ⅱ surface represents the best Zhaojue paleo-census sample giving a count of 68 individuals from 61 trackways(37 ornithopod,10 theropod,14 sauropod)and isolated tracks(representing 7 individuals).Thus,ornithopods(both large and small trackmakers)represent-54%of the total number of *** smaller ZJ-Ⅲ surface was mapped using traditional methods,and reveals at least 6 ornithopod-and 4 theropod *** the ZJ-Ⅱ and ZJ-Ⅲ surfaces reveal parallel ornithopod trackways suggestive of social or gregarious *** combined data from all four Zhaojue surfaces reveal a total of 1928 tracks,and include a few previously reported pterosaurs and theropod swim *** Zhaojue quarry complex provide a good example of multiple track-bearing sites(surfaces)that require long term study and monitoring to extract as much trackway data as possible before in situ physical evidence is lost.

关键词： Feitianshan Formation Onithopod Teropod Suropod

The Middle-Upper Jurassic dinosaur-pterosaur fauna in Shandong Province,China:Evidence from ichnology

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Journal of Palaeogeography 2023年第3期12卷 419-433页

作者： Li-Da Xing Martin G.Lockley Bo-Lin Tong Hendrik Klein Chang Liu w.scott persons IV State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing100083China Dinosaur Trackers Research Group University of Colorado DenverPO Box 173364DenverCO80217USA Saurierwelt Palaontologisches Museum Alte Richt 7D-92318NeumarktGermany Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharlestonSCUSA

Tracks previously examined between 2002 and 2015 from an incompletely-studied,and now partially lost tracksite site in the Santai Formation of Shandong Province were restudied based on new ex-cavations of additional *** contrast to the previous reports of tridactyl tracks,including Grallator,the present study discovered an additional assemblage of unequivocal pterosaur tracks(Pteraichnus)and a possible didactyl deinonychosaurian *** age of the Santai Formation is controversial but it is probably Late Jurassic,which corresponds to the epoch when pterosaur tracks appear *** interpretations,based on palynology and isotopic data,tentatively suggests an earlier Middle-Late Jurassic age and might suggest that the pterosaurian tracks are as old or somewhat older than any previously reported.

关键词： Santai Formation Theropod Deinonychosaur Pterosaur

A probable tyrannosaurid track from the Upper Cretaceous of southern China

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Science Bulletin 2019年第16期64卷 1136-1139页

作者： Lida Xing Kecheng Niu Martin G.Lockley Hendrik Klein Anthony Romilio w.scott persons IV Stephen L.Brusatte State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing 100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing 100083China Yingliang Stone Nature History Museum Nan’an 362300China Dinosaur Trackers Research Group University of ColoradoDenver 80217USA Saurierwelt Palaontologisches Museum Neumarkt D-92318Germany School of Biological Sciences the University of QueenslandQueensland 4072Australia Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA School of GeoSciences Grant InstituteUniversity of EdinburghEdinburgh EH9 3FEUK

A large number of Upper Cretaceous(Maastrichtian)dinosaur skeletal fossils have been found in the Nanxiong Basin and adjacent Ganzhou area,including those of theropods,sauropods and ***,known dinosaur ichnoassemblages from the Nanxiong Basin partly indicate a different fauna,with dominating large-and medium-sized ornithopods,theropods,and pterosaurs(1,2)This is one of many known examples where trace and body fossil assemblages from the same formation or geologically discrete region show a different *** and paleobiogeographical analyses therefore should consider data from both,before any conclusions can be *** can be classified according to the relative degree of similarity or congruity(or difference/incongruity)between trace and body fossils into the 5 categories proposed by Lockley[3].

关键词： tyrannosaurid track Upper Cretaceous southern China

New Early Cretaceous Pterosaur-Bird Track Assemblage from Xinjiang, China:Palaeoethology and Palaeoenvironment

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Acta Geologica Sinica(English Edition) 2013年第6期87卷 1477-1485页

作者： HE Qing XING Lida ZHANG Jianping Martin G.LOCKLEY Hendrik KLEIN w.scott persons IV QI Liqi JIA Chengkai School of the Earth Sciences and Resources China University of Geosciences Dinosaur Tracks Research Group University of Colorado at DenverPO Box 173364 Denver CO 8217 USA Saurierwelt Palontologisches Museum Alte Richt 7 D-92318 Neumarkt Germany Department of Biological Sciences University of Alberta 11455 Saskatchewan DriveEdmonton Alberta T6G 2E9 Canada Research Institute of Experiment and Detection Xinjiang Oilfield Company

A pterosaur-bird track assemblage from a sandstone-siltstone-mudstone sequence of the Lower Cretaceous Tugulu Group of Xinjiang comprises the first pterosaur track record from this province and the largest specimen thus far known from *** pterosaur tracks are assigned to the ichnogenus Pteraichnus based on the triangular overall-shape,the four elongate digit traces and the robust manual digit trace Ⅲ.Supposed trackmakers were dsungaripterid pterodactyloids whose skeletal remains are well known from the Tugulu *** bird tracks that occur on the same surface,are those of typical shorebirds,known from different other localities in southeast *** congruence with Koreanaornis dodsoni described from the same stratigraphic level justifies an assignment to this *** is a further evidence of the co-occurrence of pterosaurs and birds in a typical iakeshore environment with possible seasonal alteration of water supply and aerial exposure indicated by wave ripples,mudcracks and repeated cycles of coarse to fine *** and birds frequented the shoreline and may have fed also on the numerous invertebrates such as the Scoyenia tracemaker that left abundant burrows.

关键词： Pteraichnus Koreanaornis Early Cretaceous Tugulu Group Xinjiang

Unusual dinosaur trackway preservation as clues to paleo-landscape and behavior from the Lower Cretaceous Luohe Formation,Shaanxi Province,China

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Geoscience Frontiers 2021年第2期12卷 737-745页

作者： Lida Xing Martin G.Lockley Yongzhong Tang Hendrik Klein w.scott persons IV Miaoyan wang Xingwen Li Hao wu State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijingChina School of the Earth Sciences and Resources China University of GeosciencesBeijingChina Dinosaur Trackers Research Group University of Colorado DenverPO Box 173364DenverCO 80217USA Shaanxi Institute of Geological Survey Xi'an 710054ShaanxiChina Saurierwelt Paläontologisches Museum Alte Richt 7D–92318 NeumarktGermany Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA

Poorly preserved tracks have limited ichnotaxonomic or biotaxon utility,but may reveal useful information about the paleoenvironment,behavior and track *** mostly parallel to sub parallel trackway segments(T1–T8)were registered on a truncation surface in the Lower Cretaceous Luohe Formation of Shaanxi *** attest to the passage of several bipeds,probably all theropods,in a paleo-contour-parallel,south-north direction in an arid *** of preservation in these trackways is poor,but notably superior in two additional trackways(T9–T10)on a foreset *** T9 indicates a didactyl trackmaker,probably a deinonychosaurian,heading north to *** is the 13th report of deinonychosaurian tracks from the Lower Cretaceous of *** any or all the eight south-north oriented trackway segments represent continuations of other segments in the same trackways,the total number of individual trackmakers heading in this direction may have been as lowas *** the trackway pattern and sedimentological evidence could indicate a physically controlled pathway influencing the direction taken by these trackmakers,the possibility that the trackways also represented small social or gregarious group cannot be ruled out.

关键词： Theropod tracks Ornithischian tracks Deinonychosaurian Dunes sediments

The new ichnotaxon Eubrontes nobitai ichnosp. nov. and other saurischian tracks from the Lower Cretaceous of Sichuan Province and a review of Chinese Eubrontes-type tracks

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Journal of Palaeogeography 2021年第3期10卷 302-320页

作者： Li-Da Xing Martin G.Lockley Hendrik Klein Li-Jun Zhang Anthony Romilio w.scott persons IV Guang-Zhao Peng Yong Ye Miao-Yan wang State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing 100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing 100083China Dinosaur Trackers Research Group University of ColoradoDenver 80217USA Saurierwelt Paläontologisches Museum Alte Richt 7 D-92318 NeumarktGermany Institute of Resources and Environment Key Laboratory of Biogenic Traces and Sedimentary Minerals of Henan ProvinceCollaborative Innovation Center of Coalbed Methane and Shale Gas for Central Plains Economic RegionHenan Polytechnic UniversityJiaozuo 454003Henan ProvinceChina School of Biological Sciences The University of QueenslandBrisbaneQueensland 4072Australia Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA Zigong Dinosaur Museum Zigong 643013Sichuan ProvinceChina

The Jiaguan Formation and the underlying Feitianshan Formation(Lower Cretaceous) in Sichuan Province yield multiple saurischian(theropod–sauropod) dominated ichnofaunas. To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common "generic" labels such as Grallator, Eubrontes, cf. Eubrontes or even "EubrontesMegalosauripus" morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks(Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very wellpreserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.

关键词： Ichnofossils Dinosaur footprints Theropod Myths

翼龙类和驰龙类尾的前关节突和脉弧棒的趋同演化及翼龙类和鸟类尾部肌肉退化的平行模式(英文)

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地球学报 2010年第S1期31卷 54-54页

作者： w.scott persons,IV Laboratory for Vertebrate Paleontology Department of Biological SciencesUniversity of AlbertaEdmontonAlbertaCanadaT6G 2E9

The caudal morphology of dromaeosaurid dinosaurs and most "rhamphorhynchoid" pterosaurs is a striking example of convergent evolution.

关键词：翼龙类前关节突脉弧演化鸟类趋同退化肌肉尾部平行

重庆晚侏罗世两处兽脚类足迹组合与四川盆地侏罗纪地层(英文)

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古脊椎动物学报 2013年第2期51卷 107-130页

作者：邢立达 Martin G. LOCKLEY 陈伟 Gerard D.GIERLINSKI 李建军 w.scott persons Ⅳ 松川正树叶勇 Murray K.GINGRAS 王昌文中国科学院脊椎动物演化与人类起源重点实验室北京100044 美国科罗拉多大学丹佛分校恐龙足迹博物馆丹佛80217 重庆自然博物馆重庆400013 波兰侏罗纪公园奥斯特罗维茨427-4001 波兰地质研究所华沙400-975 北京自然博物馆科学研究部北京100050 加拿大阿尔伯塔大学生物科学系埃德蒙顿T6G2E9 日本东京学艺大学环境科学系东京184-8501 自贡恐龙博物馆四川自贡643013 加拿大阿尔伯塔大学地球科学系埃德蒙顿T6G2E9 重庆市永川区文物管理所重庆402160

重庆市已报道了4处侏罗纪的恐龙足迹点,包括下侏罗统珍珠冲组的大足化石点,该地点发现了中国迄今最古老的蜥脚类行迹;另外两处化石点(南岸和金鸡)的层位曾被视为中或上侏罗统,这里首次将其确认为上侏罗统上沙溪庙组;第4处化石点(成渝),... 详细信息

重庆市已报道了4处侏罗纪的恐龙足迹点,包括下侏罗统珍珠冲组的大足化石点,该地点发现了中国迄今最古老的蜥脚类行迹;另外两处化石点(南岸和金鸡)的层位曾被视为中或上侏罗统,这里首次将其确认为上侏罗统上沙溪庙组;第4处化石点(成渝), 本文全部作者目前都未曾观察,这里不做描述。南岸化石点是南岸重庆足迹(Chongqingpus nananensis)模式标本的所在地,位于重庆市中心,曾发现过大量兽脚类足迹,如今已消逝在城市化进程中。幸运的是,这批标本被保存于重庆自然博物馆,本文作者曾前后两次对其进行研究,并得出文中的结论。南岸重庆足迹为中型足迹(平均长约29 cm), 最可能被卡岩塔足迹(Kayentapus)所囊括,这批标本中有一些保存着边界不清的拇趾迹。该化石点的其他足迹被归于似异样龙足迹(cf. Anomoepus)。其他较小的、来自重庆市周边中侏罗统中部的足迹此前已被归于跷脚龙足迹(Grallator)。金鸡化石点保存了一条孤立的兽脚类行迹,因其粗壮的特征而被暂时归于似窄足龙足迹(cf. Therangospodus)。金鸡点的行迹也保存了非连续出现的拇趾迹。虽然重庆市和四川盆地及其周边更广泛地区之侏罗纪地层发现的兽脚类足迹日渐增多,但要明确其确切地质年龄及有效的足迹分类依然需要进一步工作。这是因为侏罗纪的恐龙足迹尽管丰富,但显示出连续变化的形态学特征,而保存状况的变化更使其难以在时间与空间上做出定义与鉴别。

关键词：重庆四川盆地晚侏罗世兽脚类足迹足迹分类学