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a.new species of smooth skink (Squa.a.a. Scincida.： Scincella. from Ca.bodia

Zoologica. Resea.ch 2018年第3期39卷 220-240页

作者： Thy Nea.g Soma.y Cha. nikolay a. poyarkov, Jr. Wild Ea.th a.lies Sk. Phnom Penh Thmei Kh. Sen Sok Phnom Penh Cambodia Ministry of Environment Sk. Tonle Bassac Kh. Chamkarmorn Phnom Penh Cambodia Depa.tment of Vertebra.e Zoology Biological Faculty Lomonosov Moscow State University Moscow 119234 Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Centre Nghia Do Cau Giay Hanoi Vietnam

Ba.ed on morphologica. a.d genetic evidence we eva.ua.ed the ta.onomic sta.us of a.newly discovered forest-dwelling popula.ion of skink （genus Scincella. from the Keo Selma.Wildlife Sa.ctua.y, Mondulkiri Province, Ca.bodia. From phylogenetic a.a.ysis of a.668-bp fra.ment of the mtDNa.COl a.d dia.nostic morphologica. cha.a.ters we a.loca.e the newly discovered popula.ion to the Scincella.reevesii-S. rufoca.da.a.species complex a.d describe it a. Scincella.nigrofa.cia.a.sp. nov. The new skink species ca. be distinguished from a.l other Southea.t a.ia. congeners by the following combina.ion of morphologica. cha.a.ters： snout-vent length （SVL） 40.0-52.6 mm; rela.ive ta.l length （Ta./SVL ra.io） 1.25-1.94; prefronta.s in broa. conta.t; infra.a.ia.s 6; prima.y tempora.s 2; rela.ive forelimb length （FIL/SVL ra.io） 0.20-0.22; rela.ive hindlimb length （HIL/SVL ra.io） 0.30-0.33; rela.ive forea.m length （FoL/SVL ra.io） 0.14-0.16; a.pressed forelimbs a.d hind limbs either overla.ping （0.4-2.2 mm） or sepa.a.ed （1.9-2.3 mm）; midbody sca.e rows 32-33, pa.a.ertebra. sca.es 69-74, vertebra. sca.es 65-69; dorsa. sca.es between dorsola.era. stripes 8; compa.a.ively slender fingers a.d toes, subdigita. la.ella. under fourth toe 15-17; da.k discontinuous regula. dorsa. stripes 5-7; distinct bla.k dorsola.era. stripes, na.rowing to la.era. sides a.d extending to 52%-86% of tota. ta.l length. We provide a.ditiona. informa.ion on the holotype of Scincella.rufoca.da.a.（Da.evsky ＆ Nguyen, 1983）, a.d provide evidence for the species sta.us of Scincella.rupicola. Our discovery brings the number of Scincella.pecies in Ca.bodia.to five a.d empha.izes the incompleteness of knowledge on the herpetofa.na. diversity of this country.

关键词： Mondulkiri Keo Seima.WildlifeSa.ctua.y Ta.onomy mtDNA COl DNa.ba.coding Phylogenetics

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Counting stripes:revision of the Lipinia.vittigera.complex(Reptilia. Squa.a.a. Scincida.) with description of two new species from Indochina

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Zoologica. Resea.ch 2019年第5期40卷 358-393页

作者： nikolay a. poyarkov Jr. Peter Geissler Vla.isla. a. Gorin Evgeniy a. Duna.ev Timo Ha.tma.n Cha.mongkon Suwa.na.oom Depa.tment of Vertebra.e Zoology Biological Faculty Lomonosov Moscow State University Moscow 119991 Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Center Nghia Do Cau Giay Hanoi Vietnam Museum Na.ur und Mensch Freiburg 79098 Germany Zoologica. Museum Moscow State University Moscow 125009 Russia Zoologica. Resea.ch Museum a.exa.der Koenig Bonn 53113 Germany School of a.riculture a.d Na.ura. Resources University of Phayao Phayao 56000 Thailand

We provide a. integra.ive ta.onomic a.a.ysis of the Lipinia.vittigera.species complex from ma.nla.d Southea.t a.ia. Ba.ed on exa.ina.ion of externa. morphology, color pa.tern, a.d 681 ba.e pa.rs of the cytochrome oxida.e subunit I (COI) mitochondria. gene, we demonstra.e the presence of four morphologica.ly distinct linea.es of Lipinia.in Vietna., Ca.bodia. Tha.la.d, a.d Ma.a.sia. showing a.sequence divergence ra.ging 15.5%–20.4%. a.l discovered linea.es a.e discretely dia.nosa.le from one a.other by a.combina.ion of sca.a.ion tra.ts a.d color pa.terns. a.review of the published distribution da.a.a.d a.re-exa.ina.ion of a.a.la.le type ma.eria. revea.ed the following results:(1) distribution of L. vittigera.(Boulenger, 1894) sensu stricto is restricted to Sunda.a.d a.d the Tha.-Ma.a. Peninsula.south of the Isthmus of Kra.(2) L. microcercus (Boettger, 1901) sta.. nov. is eleva.ed to full species ra.k;the species ha. a.wide distribution from centra. a.d southern Vietna. a.ross Ca.bodia.to ea.tern Tha.la.d;we rega.d Lygosoma.vittigerum kronfa.um Smith, 1922 a.d Leiolopisma.pra.ensis Cochra., 1930 a. its junior synonyms;(3) Lipinia.trivitta.a.sp. nov. occurs in hilly a.ea. of southern Vietna., Ca.bodia. a.d ea.tern Tha.la.d;a.d (4) Lipinia.va.silievi sp. nov. is currently known only from a.na.row a.ea.a.ong the Vietna.ese-Ca.bodia. border in the foothills of the centra. a.na.ite Mounta.n Ra.ge. We further provide a. identifica.ion key for Lipinia.occurring in ma.nla.d Southea.t a.ia.

关键词： Lipinia microcercus sta.. nov. Lipinia trivitta.a sp. nov. Lipinia va.silievi sp. nov. Vietna., Tha.la.d, Ca.bodia Biogeogra.hy mtDNA COI-ba.coding

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a.new species of the genus Theloderma.Tschudi, 1838 (a.phibia. a.ura. Rha.ophorida.) from Ta. Nguyen Pla.ea., centra. Vietna.

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Zoologica. Resea.ch 2018年第3期39卷 158-184页

作者： nikolay a. poyarkov Iva. I. Kropa.hev Svetla.a.S. Gogoleva.Nikola. L. Orlov Depa.tment of Vertebra.e Zoology Biological Faculty Lomonosov Moscow State University Moscow 119234 Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Center Nghia Do Cau Giay Hanoi Vietnam Tula.Exota.ium Tula 300004 Russia a.N. Severtsov Institute of Ecology a.d Evolution Russian Academy of Sciences Moscow 119071 Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Center South Branch Ho Chi Minh City Vietnam Zoologica. Museum of the Lomonosov Moscow Sta.e University Moscow 125009 Russia Zoologica. Institute Russian Academy of Sciences St. Petersburg 199034 Russia

a.new species of sma.l tree frog from a.prima.y monta.e tropica. forest of centra. Vietna., Ta. Nguyen Pla.ea., is described ba.ed on morphologica., molecula., a.d a.oustic evidence. The Golden Bug-Eyed Frog, Theloderma.a.ra.um sp. nov., is distinguisha.le from its congeners a.d other sma.l rha.ophorid species ba.ed on a.combina.ion of the following morphologica. a.tributes：（1） bony ridges on hea. a.sent; （2） smooth skin completely la.king ca.cified wa.ts or a.perities; （3） pointed elonga.ed ta.ering snout; （4） voca. opening in ma.es a.sent; （5） vomerine teeth a.sent; （6） ma.es of sma.l body size （SVL 21.8-26.4 mm）; （7） hea. longer tha. wide; ED/SVL ra.io 13%-15%; ESL/SVL ra.io 16%-20%; （8） sma.l tympa.um （TD/EL ra.io 50%-60%） with few tiny tubercles; （9） supra.ympa.ic fold a.sent; （10） ventra. surfa.es completely smooth; （11 ） webbing between fingers a.sent; （12） outer a.d inner meta.a.pa. tubercles present, supernumera.y meta.a.pa. tubercle single, media., ova. in sha.e; （13） toes ha.f-webbed： I 2-21/4 II 11/^-23/~ III 2-31/4 IV 3-11/2 V; （14） inner meta.a.sa. tubercle present, ova.; outer meta.a.sa. tubercle a.sent; （15） iris bicolored; （16） dorsa. surfa.es golden-yellow with spa.se golden-ora.ge speckling or reticula.ions a.d few sma.l da.k-brown spots; （17） la.era. sides of hea. a.d body with wide da.k reddish-brown to bla.k la.era. stripes, clea.lysepa.a.ed from lighter dorsa. colora.ion by stra.ght contra.ting edge; （18） ventra. surfa.es of body, throa., a.d chest greyish-blue with indistinct brown confluent blotches; （19） upper eyelids with few （3-5） very sma.l fla. reddish supercilia.y tubercles; （20） limbs dorsa.ly reddish-brown, ventra.ly brown with sma.l bluish-white speckles. The new species is a.so distinct from a.l congeners in 12S rRNa.to 16S rRNa.mitochondria. DNa.fra.ment sequences （uncorrected genetic dista.ce P〉8.9%）. a.vertisement ca.l a.d ta.pole morphology of the new species a.e described. Our molecula. da.a.showed Theloderma.a.ra.um sp. nov.

关键词： Theloderma.a.ra.um sp. nov. mtDNa.hylogeny 12S rRNA 16S rRNA Kon Tum Gia.La. Endemism Ta.onomy

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a.new species of the genus Leptola.a. (a.ura. Megophryida.) from southern Vietna.

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Zoologica. Resea.ch 2018年第3期39卷 185-201页

作者： Ta.g Va. Duong Da.g Trong Do Chung Da. Ngo Truong Qua.g Nguyen nikolay a. poyarkov Vietna. Na.iona. Museum of Na.ure Vietnam Academy of Science and Technology Hanoi Vietnam Depa.tment of Vertebra.e Zoology Biological FacultyLomonosov Moscow State University Moscow 119234 Russia Fa.ulty of Na.ura. Sciences Phu Yen University Tuy Hoa Phu Yen Vietnam College of Educa.ion Hue University Hue Vietnam Institute of Ecology a.d Biologica. Resources Vietnam Academy of Science and Technology Hanoi Vietnam Gra.ua.e University of Science a.d Technology Vietnam Academy of Science and Technology Cau Giay Hanoi Vietnam Joint Russia.- Vietna.ese Tropica. Resea.ch a.d Technologica. Center Nghia Do Cau Giay Hanoi Vietnam

We describe a.new species of megophryid frog from Phu Yen Province in southern Vietna.. Leptola.a. ma.rops sp. nov. is distinguished from its congeners by a.combina.ion of the following morphologica. a.tributes：（1） body size medium （SVL 28.0-29.3 mm in three a.ult ma.es, 30.3 mm in single a.ult fema.e）; （2） supra.a.illa.y gla.ds present, crea.y white; ventrola.era. gla.ds indistinct; （3） tympa.um externa.ly distinct; （4） dorsa. skin roughly gra.ula. with la.ger tubercles, derma. ridges on dorsum a.sent; （5） rudimenta.y webbing present between fingers I-II a.d I1-111; rudimenta.y webbing between a.l toes; fingers a.d toes without derma. fringes; （6） in life ventra. surfa.e greyish-violet with white speckling; （7） supra.ympa.ic fold distinct, da.k brown in life; （8） iris bicolored, typica.ly golden in upper ha.f, fa.ing to golden green in lower ha.f; （9） tibia.short （TbL/SVL 0.44-0.45 in ma.es）; a.d （10） eyes la.ge a.d protubera.t （ED/SVL 0.15-0.16 in ma.es） From a.l congeners for which compa.a.le sequences a.e a.a.la.le, the new species differs ma.kedly in the 16S rRNa.mitochondria. gene sequence （P-dista.ce〉5.7%）. The new species is currently known only from monta.e evergreen tropica. forests of Song Hinh District, Phu Yen Province, a.d M＇Dra. District of Da. La. Province a. eleva.ions of 470-630 m a.s.l.. We suggest the new species should be considered a. Da.a.Deficient following the IUCN＇s Red List ca.egories. We a.so report a.previously unknownLeptola.a. mtDNa.linea.e from a. evergreen tropica.forest in the Hoa.Thinh District of Phu Yen Province which ma. a.so represent a. undescribed species.

关键词： Leptola.a. ma.rops sp. nov. Phu YenProvince Da. La. Province Southern coa.ta. regionof Vietna.

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On the roa. to Ma.da.a.: contribution to the Microhyla.Tschudi, 1838(a.phibia. a.ura. Microhylida.) fa.na.of Mya.ma. with description of two new species

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Zoologica. Resea.ch 2019年第4期40卷 244-276页

作者： nikolay a. poyarkov, Jr Vla.isla. a. Gorin Tha. Za. Va.entina.D. Kretova.Svetla.a.S. Gogoleva.Pa.inya.Pa.a.gkha.a.t Jing Che Depa.tment of Vertebra.e Zoology Biological FacultyLomonosov Moscow State UniversityMoscow 119234Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Center Nghia DoCau GiayHanoiVietnam Zoology Depa.tment Mohnyin Degree CollegeMohnyinKachin State 1111Myanmar a.N. Severtsov Institute of Ecology a.d Evolution Russian Academy of SciencesMoscow 119071Russia Zoologica. Museum of the Lomonosov Moscow Sta.e University Moscow 125009Russia Ba.somdejcha.pra.a.Ra.a.ha. University Thon BuriBangkok 10600Thailand Sta.e Key La.ora.ory of Genetic Resources a.d Evolution Kunming Institute of ZoologyChinese Academy of SciencesKunming Yunnan 650223China Southea.t a.ia.Biodiversity Resea.ch Institute Chinese Academy of SciencesYezin Nay Pyi Taw 05282Myanmar

We prese nt a.morphologica. a.d molecula. a.sessment of the Microhyla.fa.na.of Mya.ma. ba.ed on new collections from centra. (Ma.wa. Division) a.d northem (Ka.hin Sta.e) pa.ts of the country. In tota., six species of Microhyla.a.e documented, including M. berdmorei, M. heymonsi, M. butleri, M. mukhlesuri a.d two new species described from the semi-a.id sa.a.ma.like pla.ns of the middle pa.t of the Irra.a.dy (a.eya.wa.y) River Va.ley. We used a.2 481 bp long 12S rRNa. 16S rRNa.fra.ment of mtDNa.to hypothesize gen ea.ogica. rela.i on ships within Microhyla. We a.plied a. in teg ra.i ve ta.onomic a.proa.h combining molecula., morphologica., a.d a.oustic lines of evidenee to eva.ua.e the ta.onomic sta.us of Mya.nma. Microhyla. We dem on stra.ed tha. the newly discovered popula.ions of Microhyla.sp. from the Ma.wa. Division represent two yet undescribed species. These two new sympa.ric species a.e a.signed to the M. a.ha.ina.species group, with both a.a.ted to the sea. on a.ly dry environments of the Irra.a.dy Va.ley. Microhyla.fodiens sp. nov. is a.stout-bodied species with a.rema.ka.ly enla.ged shovel-like outer meta.a.sa. tubercle used for burrowing a.d is highly diverge nt from other know n congeners (P-dista.cea..8%). Microhyla.irra.a.dy sp. nov. is a.sma.l-bodied slender frog reconstructed a. a.sister species to M. kodia. from southern India.(P-dista.ce=5.3%);however, it clea.ly differs from the la.ter both in externa. morphology a.d a.vertisement ca.l pa.a.eters. Microhyla.mukhlesuri is reported from Mya.ma. for the first time. We further discuss the morphologica. dia. no sties a.d biogeogra.hy of Microhyla.species recorded in Mya.ma..

关键词： Na.row-mouth frogs Burma Indochina Ma.wa. Ka.hin Biodiversity Ta.onomy mtDNA Morphology a.oustics a.vertisement ca.l

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a.new ka.st-dwelling bent-toed gecko (Squa.a.a. Gekkonida. Cyrtoda.tylus) from Xia.gkhoa.g Province, northea.tern La.s

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Zoologica. Resea.ch 2018年第3期39卷 202-219页

作者： Roma. a. Na.a.ov Olivier S.G. Pa.wels Evgeniy L. Konsta.tinov a.a.oliy S. Chulisov Nikola. L. Orlov nikolay a. poyarkov, Jr. Zoologica. Museum Moscow State University Moscow 125009 Russia Roya. Belgia. Institute of Na.ura. Sciences Brussels B-1000 Belgium Institute of Na.ura. Sciences Kaluga State University named after K.I. Tsiolkovskii Kaluga 248023 Russia Zoologica. Institute Russian Academy of Sciences St. Petersburg 199034 Russia Depa.tment of Vertebra.e Zoology Biological Faculty Lomonosov Moscow State University Moscow 119234 Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Center 63 Nguyen Van Huyen Road Nghia Do Cau Giay Hanoi Vietnam

We describe a.new ka.st-dwelling Cyrtoda.tylus from Ba. Tha.hom, Xia.gkhoa.g Province, northea.tern La.s. The new species ca. be distinguished from other congeners by ha.ing four da.k dorsa. ba.ds between limb insertions, a.discontinuous nucha. loop, 10 precloa.a. pores in ma.es or 10-12 precloa.a. pits （fema.es） sepa.a.ed by a.dia.tema.from a.series of enla.ged femora. sca.es bea.ing 18 or 19 pores （ma.e） or 8-10 pits （fema.es） a.ong ea.h femur, 14-18 dorsa. tubercle rows a. midbody, no precloa.a. groove, 30-36 midbody sca.e rows a.ross belly between ventrola.era. skin folds, tra.sversely enla.ged subca.da. pla.es, a.d a.ma.ima. known snout-vent length of 75.5 mm. Our description brings to 22 the number of Cyrtoda.tylus species recorded from La.s.

关键词： Indochinese region Ka.st Limestone Herpetology Ta.onomy New species

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a.new species of the genus Xenophrys(a.ura.Megophryida.) from northern Tha.la.d

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Zoologica. Resea.ch 2019年第6期40卷 564-574页

作者： Yun-He Wu Cha.mongkon Suwa.na.oom nikolay a. poyarkov,Jr. Pa.inya.Pa.wa.gkha.a.t Ka. Xu Jie-Qiong Jin Robert W. Murphy Jing Che Sta.e Key La.ora.ory of Genetic Resources a.d Evolution Sta.e Kunming Institute of ZoologyChinese Academy of SciencesKunming Yunnan 650223China Kunming College of Life Science University of the Chinese Academy of SciencesKunming Yunnan 650204China Southea.t a.ia.Biodiversity Resea.ch Institute Chinese Academy of SciencesYezin Nay Pyi Taw 05282Myanmar School of a.riculture a.d Na.ura. Resources University of PhayaoPhayao 56000Thailand Depa.tment of Vertebra.e Zoology Biological FacultyLomonosov Moscow State UniversityMoscow 119991Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Center 63 Nguyen Van Huyen RoadNghia DoCau GiayHanoiVietnam Centre for Biodiversity a.d Conserva.ion Biology Royal Ontario MuseumToronto M5S 2C6Canada

DEa. EDITOR,Species of Xenophrys a.e conserved morphologica.ly a.d live prima.ily in forests. In Tha.la.d, the genus ha.bors ma.y cryptic species. Herein we report the collection of specimens from Doi Intha.on, Chia.g Ma. Province, northern Tha.la.d, which were identified previously a. X. minor. Molecula. a.d morphologica. a.a.yses find tha. these specimens differ significa.tly from other known congeners, a.d therefore we describe a.new species. Further, our phylogenetic a.a.yses indica.e tha. X. la.ida.tyla.is a.junior synonym of X. pa.pebra.espinosa.

关键词： genus Xenophrys Megophryida. Tha.la.d

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a.new species of the genus Micryletta.a.ura.Microhylida.) from Ha.na. Isla.d, China

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Zoologica. Resea.ch 2021年第2期42卷 234-240页

作者： Jia.-Hua. Ya.g nikolay a.poyarkov Ka.oorie Conserva.ion China Kadoorie Farm and Botanic GardenTai PoHong Kong SARChina Fa.ulty of Biology Department of Vertebrate ZoologyMoscow State UniversityMoscow 119234Russia La.ora.ory of Tropica. Ecology Joint Russian-Vietnamese Tropical Research and Technological CenterHanoiVietnam

DEa. EDITOR,We report on a.new a.phibia. species of the genus Micryletta.from Ha.na. Isla.d, China. ba.ed on morphologica. a.d molecula. a.a.yses. The new species, Micryletta.imma.ula.a.sp. nov., is dia.nosed by a.combina.ion of the following morphologica. cha.a.ters: medium-sized within genus(SVL23.3–24. mm in ma.es, n=3;27.7–30.1 mm in fema.es, n=2).

关键词： nov. Isla.d new

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a.new species of Micryletta.a.phibia.a.ura.Microhylida.)from the La.gbia. Pla.ea. in southern Vietna.

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Zoologica. Resea.ch 2021年第6期42卷 726-733页

作者： nikolay a.poyarkov Ta. Va. Nguyen Jia.-Hua. Ya.g Vla.isla. a.orin Fa.ulty of Biology Department of Vertebrate ZoologyMoscow State UniversityMoscow119234Russia La.ora.ory of Tropica. Ecology Joint Russian-Vietnamese Tropical Research and Technological CenterHanoiVietnam Depa.tment of Species Conserva.ion Save Vietnam’s WildlifeNinh BinhVietnam Ka.oorie Conserva.ion China Kadoorie Farm and Botanic GardenTai PoHong Kong SARChina

We report a.new species of the genus Micryletta.from the monta.e evergreen forest in the Bidoup-Nui Ba.Na.iona. Pa.k,La. Dong Province,La.gbia. Pla.ea.,southern Vietna.,ba.ed on molecula. a.d morphologica. *** new species is dia.nosed by a.combina.ion of the following morphologica. cha.a.ters:body size sma.l(snout-vent length(SVL)22.4 mm,single fema.e);iris uniform bla.k;snout nea.ly trunca.e in dorsa. view,slightly rounded in profile;tibiota.sa. a.ticula.ion of a.pressed limb rea.hing level of eye;dorsa. surfa.e smooth;supra.ympa.ic fold present,prominent;outer meta.a.sa. tubercle a.sent;tips of toes very wea.ly dila.ed into sma.l discs;finger webbing a.sent,toe webbing rudimenta.y;dorsa. surfa.es of hea. a.d body ora.ge-red with sma.l irregula.ly sha.ed da.k-brown pa.ches;dorsa. surfa.e of limbs pa.e da.k brown with sma.l reddish speckles;body fla.ks da.k brown a.teriorly fa.ing to gra.ish brown posteriorly with brown spots in groin a.ea.la.era. sides of hea. imma.ula.e bla.kish brown without white pa.ches;colora.ion of ventra. surfa.es imma.ula.e da.k gra..

关键词： Pla.ea. ora.ge brown

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a.new species of Crocodile Newt, genus Tylototriton(a.phibia. Ca.da.a. Sa.a.a.drida.) from the mounta.ns of Ka.hin Sta.e, northern Mya.ma.

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Zoologica. Resea.ch 2019年第3期40卷 151-174页

作者： Tha. Za. Pa. La. Pa.inya.Pa.a.gkha.a.t Vla.isla. a.Gorin nikolay a.poyarkov,Jr Zoology Depa.tment Mohnyin Degree CollegeMohnyinKachin StateMyanmar Flora.a.d Fa.na.Interna.iona. Lon Ton VillageIndawgyiKachin StateMyanmar Ba.somdejcha.pra.a.Ra.a.ha. University Hiran RuchiThon BuriBangkok 10600Thailand Depa.tment of Vertebra.e Zoology Biological FacultyLomonosov Moscow State UniversityMoscow 119234Russia Joint Russia.-Vietna.ese Tropica. Resea.ch a.d Technologica. Center Nghia DoCau GiayHanoiVietnam

We describe a.new species of the genus Tylototriton from Ingyin Ta.ng Mt.,Mohnyin Township,Ka.hin Sta.e,Mya.ma.,ba.ed on morphologica. a.d molecula. *** new species is a.signed to the subgenus Tylototriton *** is clea.ly distinct from a.l known congeners by the following cha.a.ters:medium body size;thin,long ta.l,la.king la.era. grooves;rough skin;trunca.e snout;wide,protruding supra.empora. bony ridges on hea.,beginning a. a.terior corner of orbit;wea.,a.most indistinct sa.itta. ridge;long,thin limbs,broa.ly overla.ping when a.pressed a.ong body;distinct,wide,non-segmented vertebra. ridge;13 or 14 rib nodules;brown to da.kbrown ba.kground colora.ion with dull ora.ge-brown to yellowish-brown ma.kings on la.ia. regions,pa.otoids,rib nodules,whole limbs,vent,a.d ventra. ta.l *** a.so briefly discuss biogeogra.hy a.d species diversity of the genus Tylototriton in Mya.ma..

关键词： Tylototriton ka.hinorum sp.nov. mtDNA genea.ogy ND2 16S rRNA Sha. Biogeogra.hy Endemism Ta.onomy

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