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Study on fault-controlled hydrocarbon migration and accumulation process and models in Zhu I Depression

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Acta Oceanologica Sinica 2021年第2期40卷 107-113页

作者： Wenqi Zhu keqiang wu Ling Ke Kai Chen Zhifeng Liu CNOOC Research Institute Co. Ltd.Beijing 100028China

Through the analysis of the faults and their internal structure in Zhu I Depression,it is found that the internal structure of the late fault is obviously segmented *** develops unitary structure(simple fault plane)in shallow layers,binary structure(induced fracture zone in hanging wall and sliding fracture zone in footwall)in middle,layers and ternary structure(induced fracture zone in hanging wall and sliding fracture zone in middle,and induced fracture zone in footwall)in deep *** the induced fracture zone is a high porosity and permeability zone,and the sliding fracture zone is a low porosity and ultra-low permeability zone,the late fault in middle layers has the character of"transporting while sealing".The late fault can transport hydrocarbon by its induced fracture zone in the side of the hanging wall and seal hydrocarbon by its sliding fracture zone in the side of the *** deep layers,the late fault has the character of"dual-transportation",induced fracture zones in both sides of hanging wall and footwall can transport *** early fault that only developed in the deep layers is presumed to be unitary structure,which plays a completely sealing role in the process of hydrocarbon migration and accumulation due to inactivity during the hydrocarbon filling *** by hydrocarbon source,early/late faults,sand bodies and traps,two reservoir-forming models of"inverted L"and"stereo-spiral"can be proposed in middle layers,while two reservoir-forming models of"cross fault"and"lateral fault sealing"are developed in the deep layers of Zhu I Depression.

关键词： fault structure transport/sealing migration and accumulation process reservoir-forming model Zhu I Depression

Involvement of Histone Modifcations in Plant Abiotic Stress Responses

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Journal of Integrative Plant Biology 2013年第10期55卷 892-901页

作者： Lianyu Yuan Xuncheng Liu Ming Luo Songguang Yang keqiang wu Key Laboratory of Plant Resources Conservation and Sustainable Utilization South China Botanical Garden the Chinese Academy of Sciences University of Chinese Academy of Sciences Institute of Plant Biology College of Life Science National Taiwan University

As sessile organisms, plants encounter various environmental stimuli including abiotic stresses during their lifecycle. To survive under adverse conditions, plants have evolved intricate mechanisms to perceive external signals and respond accordingly. Responses to various stresses largely depend on the plant capacity to modulate the transcriptome rapidly and specifically. A number of studies have shown that the molecular mechanisms driving the responses of plants to environmental stresses often depend on nucleosome histone post-translational modifications including histone acetylation, methylation, ubiquitination, and phosphorylation. The combined effects of these modifications play an essential role in the regulation of stress responsive gene expression. In this review, we highlight our current understanding of the epigenetic mechanisms of histone modifications and their roles in plant ahiotic stress response.

关键词： Abiotic stresses gene regulation histone modifications,

Dissolution mechanism of a deep-buried sandstone reservoir in a deep water area:A case study from Baiyun Sag,Zhujiang River(Pearl River)Mouth Basin

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Acta Oceanologica Sinica 2023年第3期42卷 151-166页

作者： Jihua Liao keqiang wu Lianqiao Xiong Jingzhou Zhao Xin Li Chunyu Zhang China National Offshore Oil Corporation(CNOOC)Research Institute Co. Ltd.Beijing 100028China Hainan Branch of China National Offshore Oil Corporation(CNOOC)Co. Ltd.Haikou 570311China School of Earth Sciences and Engineering Xi’an Shiyou University Xi’an Petroleum UniversityXi’an 710065China

Dissolution mechanism and favorable reservoir distribution prediction are the key problems restricting oil and gas exploration in deep-buried *** this paper,the Enping Formation and Zhuhai Formation in Baiyun Sag of South China Sea was taken as a *** on the thin section,scanning electron microscopy,X-ray diffraction,porosity/permeability measurement,and mercury injection,influencing factors of dissolution were examined,and a dissolution model was ***,high-quality reservoirs were predicted temporally and *** results show that dissolved pores constituted the main space of the Paleogene sandstone *** primarily occurred in the coarse-and medium-grained sandstones in the subaerial and subaqueous distributary channels,while dissolution was limited in fine-grained sandstones and inequigranular *** main dissolved minerals were feldspar,tuffaceous matrix,and diagenetic *** of feldspar and illitization of kaolinite are the main dissolution pathways,but they occur at various depths and temperatures with different geothermal *** is controlled by four factors,in terms of depositional facies,source rock evolution,overpressure,and fault activities,which co-acted at the period of 23.8–13.8 Ma,and resulted into strong ***,based on these factors,high-quality reservoirs of the Enping and Zhuhai formations are predicted in the northern slope,southwestern step zone,and Liuhua uplift in the Baiyun Sag.

关键词： dissolution mechanism deep-buried reservoir diagenesis evolution reservoir prediction deep water region Baiyun Sag

Arabidopsis NF-YCs Mediate the Light-Controlled Hypocotyl Elongation via Modulating Histone Acetylation

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Molecular Plant 2017年第2期10卷 260-273页

作者： Yang Tang Xuncheng Liu Xu Liu Yuge Li keqiang wu Xingliang Hou Key Laboratory of South China Agricultural Plant Molecular Analysis and Genetic Improvement & Guangdong Provincial Key Laboratory of Applied Botany South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China University of the Chinese Academy of Sciences Beijing 100049 China Institute of Plant Biology College of Ufe Science National Taiwan University Taipei 106 Taiwan These authors contributed equally to this article.

Light is a crucial environmental signal that promotes photomorphogenesis, the developmental process with a series of light-dependent alterations for plants to adapt various external challenges. Chromatin modification has been proposed to be involved in such light-mediated growth, but the underlying mecha- nism is still elusive. In this study, we identified four Arabidopsis thaliana Nuclear Factor-YC homologs, NF- YCl, NF-YC3, NF-YC4, and NF-YC9 （NF-YCs）, which function redundantly as repressors of light-controlled hypocotyl elongation via histone deacetylation. Obvious etiolation phenotypes are observed in NF-YCs loss-of-function mutant seedlings grown under light conditions, including significant elongated hypocotyls and fewer opened cotyledons. We found that NF-YCs interact with histone deacetylase HDA15 in the light, co-target the promoters of a set of hypocotyl elongation-related genes, and modulate the levels of histone H4 acetylation on the associated chromatins, thus repressing gene expression. In contrast, NF-YC-HDA15 complex is dismissed from the target genes in the dark, resulting in increased level of H4 acetylation and consequent etiolated growth. Further analyses revealed that transcriptional repression activity of NF-YCs on the light-controlled hypocotyl elongation partially depends on the deacetylation activity of HDA15, and loss of HDA15 function could rescue the short-hypocotyl phenotype of NF-YCs overexpression plants. Taken together, our results indicate that NF-YCl, NF-YC3, NF-YC4, and NF-YC9 function as tran- scriptional co-repressors by interacting with HDA15 to inhibit hypocotyl elongation in photomorphogen- esis during the early seedling stage. Our findings highlight that NF-YCs can modulate plant development in response to environmental cues via epigenetic regulation.

关键词： photomorphogenesis hypocotyl elongation NF-YC histone deactylation

Reassessment of the Distribution of Mantle CO_(2) in the Bohai Sea,China:The Perspective from the Source and Pathway System

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Acta Geologica Sinica(English Edition) 2022年第1期96卷 337-347页

作者： TANG Huafeng GUO Tianchan wu keqiang LIU Zilin XU Jianyong LU Baoliang WANG Pujun Key Laboratory for Evolution of Past Life and Environment in Northeast Asia Jilin UniversityChangchun 130061China Key Laboratory of Mineral Resources Evaluation in Northeast Asia Ministry of Natural ResourcesChangchun 130061China College of Earth Sciences Jilin UniversityChangchun 130061China CNOOC Research Institute Beijing 100028China Institute of Gravity and Magnetic Technology Chang’an UniversityXi’an 710054China

Research on the distribution of mantle CO_(2)should involve comprehensive analysis from CO_(2)source to *** crust-mantle pathway system is the key controlling factor of the distribution of mantle CO_(2),but has received little *** pathway system and controlling factors of CO_(2)distribution in the Bohai Sea are analyzed using data on fault styles and information on the mantle and *** relation between volcanic rocks and the distribution of mantle CO_(2)is reassessed using age data for CO_(2)*** distribution of mantle CO_(2)is controlled by uplift of the asthenosphere and upper mantle,magma conduits in the mantle and fault systems in the *** regions of the asthenosphere are accumulation areas for CO_(2).The area with uplift of the Moho exhibits accumulation of mantle CO_(2)at ***_(2)was mainly derived from vertical migration through the upper mantle and lower *** fault style in the upper crust controls the distance of horizontal migration and the locations of CO_(2)*** distribution of mantle CO_(2)and volcanic rocks are not the same,but both probably followed the same pathways *** CO_(2)in the Bohai Sea is concentrated in the Bozhong sag and the surrounding area,particularly in a trap that formed before 5.1 Ma and is connected to crustal faults(the Bozhang faults)and lithospheric faults(the Tanlu faults).

关键词： mantle CO_(2) source area pathway system fault Moho Bohai Sea

WHIRLY1 recruits the histone deacetylase HDA15 repressing leaf senescence and flowering in Arabidopsis

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Journal of Integrative Plant Biology 2022年第7期64卷 1411-1429页

作者： Dongmei Huang Wei Lan Weibo Ma Rulin Huang Wenfang Lin Mengsi Li Chia-Yang Chen keqiang wu Ying Miao Fujian Provincial Key Laboratory of Plant Functional Biology College of Life SciencesFujian Agriculture and Forestry UniversityFuzhou 350002China Institute of Botany College of Life SciencesTaiwan UniversityTaibei 106China

Leaf senescence is controlled by a complex regulatory network in which robustness is ensured by the activity of transcription factors and epigenetic ***,how these coordinate the process of leaf senescence remains poorly *** found that WHIRLY1 interacts with Histone Deacetylase(HDA)15,a Reduced Potassium Dependence3(RPD3)/HDA1-type HDA,by using green fluorescent protein-nanotrap-mass spectrum *** development-dependent interaction between WHIRLY1 and HDA15 was further confirmed by bimolecular fluorescence complementation assays and co-immunoprecipitation assays in ***-omics genome-wide transcriptome and H3K9 acetylome enrichment analysis showed that HDA15 delays leaf senescence and flowering by repressing the expression of the positive regulators of leaf senescence and flowering,such as LOX2 and LARP1 C,and reducing H3K9 ac levels at these loci;WHIRLY1 and HDA15 co-target to the region near the transcription start site of a subset of nutrient recycling-related genes(e.g.,Glutathione S-transferases 10,non-coding RNA,and photosystem II protein D1 synthesizer attenuator PDIL1-2),as well as WRKY53 and ELF4,and co-repress their expression by removing H3K9 *** study revealed a key transcription regulatory node of nutrient recycling and senescence-associated genes involved in leaf senescence and flowering via the recruitment of HDA15 by the single-stranded DNA/RNA-binding protein WHIRLY1.

关键词： flowering HDA15 H3K9 acetylation leaf senescence multi-omics analysis WHY1

The Expression of Manganese Superoxide Dismutase Gene from Nelumbo nucifera Responds Strongly to Chilling and Oxidative Stresses

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Journal of Integrative Plant Biology 2009年第3期51卷 279-286页

作者： Wen Li Lin Qi Xiaodong Lin Huhui Chen Zhanqi Ma keqiang wu Shangzhi Huang School of Life Sciences Sun Yat-sen University Guangzhou 510275 China Institute of Plant Biology School of Life Sciences National Taiwan University Taipei 106 China

A manganese superoxide dismutase （Mn-SOD） gene, NnMSD1, was identified from embryonic axes of the sacred lotus （Nelumbo nucifera Gaertn.）. The NnMSD1 protein contains all conserved residues of the Mn-SOD protein family, including four consensus metal binding domains and a signal peptide for mitochondrial targeting. Southern blot analysis suggests the existence of two *** genes in sacred lotus. NnMSD1 was highly expressed in developing embryonic axes during seed development, but appeared in cotyledons only at the early stage of development and became undetectable in the cotyledons during late embryogenesis. The expression of the NnMSD1 gene in germinating embryonic axes, in response to various stresses such as heat shock, chilling, and exposure to stress-related chemicals, was also studied. Heat shock strongly inhibited the expression of the NnMSD1 gene, whereas the NnMSD1 transcript level increased strongly in chilling stress treatment. An increase in expression was also highly induced by H2O2 in germinating embryonic axes. The results suggest that the expression pattern of the NnMSD1 gene differed between developing axes and cotyledons, and that the NnMSD1 gene expression responds strongly to chilling and oxidative stress.

关键词： manganese superoxide dismutase sacred lotus (Nelumbo nucifera Gaertn.) seed development and germination stress.

Expression and subcellular localization of histone deacetylases in tomato(Solanum lycopersicum)

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Expression and subcellular localization of histone deacetyla...

珠江口盆地白云凹陷深层储层特征与有效储层控制因素

2013全国植物生物学大会

作者： LU Jingxia YANG Songguang wu keqiang South China Botanical Garden Chinese Academy of Sciences

Recent studies have indicated that the histone acetylation plays an important role in plant development and plant response to environmental *** tomato(Solanum lycopersicum),the respiratory

关键词： histone deacetylases tomato (Solanum lycopersicum) gene expression subcellular localization

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地球科学 2022年第7期47卷 2454-2467页

作者：廖计华吴克强耳闯中海油研究总院有限责任公司北京100028 西安石油大学地球科学与工程学院陕西西安710065

珠江口盆地白云凹陷珠海组和恩平组是该区深层油气勘探的主要层系,查明深层储层基本特征和有效储层控制因素对于白云凹陷深层油气勘探具有重要意义.基于岩石学和矿物学、成岩作用、孔隙特征及沉积相分析,明确了白云凹陷深层储层基本特征... 详细信息

珠江口盆地白云凹陷珠海组和恩平组是该区深层油气勘探的主要层系,查明深层储层基本特征和有效储层控制因素对于白云凹陷深层油气勘探具有重要意义.基于岩石学和矿物学、成岩作用、孔隙特征及沉积相分析,明确了白云凹陷深层储层基本特征.白云凹陷深层以低孔低渗和致密储层为主;压实作用是造成深层储层变差的主要原因,碳酸盐胶结和石英次生加大是主要的自生矿物;孔隙类型以粒间溶孔和粒内溶孔为主;有效储层以低孔低渗及以上储层为主,孔隙度总体保持在10%左右,渗透率变化范围大.中粗粒沉积相带、溶蚀作用和超压是深层有效储层的主要控制因素.中粗粒砂岩具有较好的原生孔和次生孔发育条件,渗透率较高,胶结减孔作用弱,溶蚀增孔作用强;溶蚀作用是深层关键的建设性成岩作用,溶蚀孔隙是深层主要的孔隙类型;超压传导作用有利于酸性流体活动和溶蚀物质的迁移,对形成溶蚀孔隙具有积极意义;分流河道和水下分流河道砂体是中粗粒砂岩的主要载体,应作为深层油气勘探的优选对象.

关键词：深层储层中粗粒相带溶蚀作用超压白云凹陷石油地质学