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Geoscience Frontiers 2018年第6期9卷 1745-1754页

作者： Lida Xing Martin G.Lockley hendrik klein Rong Zeng Sifu Cai Xiuchun Luo Chen Li State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing 100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing 100083China State Key Laboratory of Palaeobiology and Stratigraphy Nanjing Institute of Geology and PalaeontologyChinese Academy of SciencesNanjing 210008China Dinosaur Tracks Museum University of Colorado DenverPO Box 173364DenverCO 80217USA Saurierwelt Palantologisches Museum Alte Richt 7D-92318 NeumarktGermany Guizhou Provincial Museum Guiyang 550001China

A newly discovered Jiaguan Formation(Lower Cretaceous) tracksite from the Linjiang region of Guizhou Province, China, reveals the first example of a Cretaceous track morphotype attributable to the non-avian theropod ichnogenus Gigandipus, here named Gigandipus chiappei ichnosp nov. The theropod dominated locality also reveals the second report of the avian theropod ichnogenus Wupus, one of the largest avian traces currently known from the Lower Cretaceous. The Linjiang site provides evidence to support previous interpretations of a distinctive Lower Cretaceous theropod-dominated ichnofauna that was widespread in China and East Asia and highlights the similarity between Lower Cretaceous theropod ichnotaxa in East Asia and those found in the Lower Jurassic both in East Asia and elsewhere. These similarities in turn create various ichnotaxonomic challenges familiar to researchers working on theropod tracks, and we recommend caution in the naming of new theropod ichnotaxa at the ichnogenus level.

关键词： Early Cretaceous Jiaguan formation Theropod tracks Gigandipus Wupus

Early Jurassic basal sauropodomorpha dominated tracks from Guizhou, China: Morphology, ethology, and paleoenvironment

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Geoscience Frontiers 2019年第1期10卷 229-240页

作者： Lida Xing Martin G. Lockley Dongjie Tang hendrik klein Guangzhao Peng Yong Ye Baoqiao Hao State Key Laboratory of Biogeology and Environmental Geology China University of Geosciences School of Earth Sciences and Resources China University of Geosciences State Key Laboratory of Palaeobiology and Stratigraphy Nanjing Institute of Geology and Palaeontology Chinese Academy of Sciences Dinosaur Trackers Research Group CB 172 University of Colorado at Denver Saurierwelt Pal?ontologisches Museum Zigong Dinosaur Museum

The newly discovered large(350 m^2) Yantan dinosaur tracksite, in the Lower Jurassic Ziliujing Formation of Guizhou Province, China, reveals at least 250 footprints of which ~97 can be resolved into trackways of sauropodomorphs. All the trackways are sub parallel likely indicating gregarious behavior. One theropod track(cf. Grallator) was recorded. The sauropodomorph tracks predominantly represent quadrupedal progression(Morphotype A), and footprint morphology is similar to the ichnospecies Liujianpusshunan, characterized by outward pes rotation. Three trackways indicate bipedal progression, and two of these(Morphotype B) indicate inward pes rotation, accompanied by elongate pes digit scratch marks. For the latter phenomenon three possible scenarios are discussed:(1) significant rotation changes accompanying changes in gait,(2) swimming behavior,(3) formation of *** evidence indicates the tracks were made on a linguloid rippled, muddy, immature sandstone substrate characterized by significant differences in substrate consistency across the trackbearing surface. Microbially induced sedimentary structures(MISS) characterized by distinctive wrinkle marks indicate a stressed, probably semi-arid, paleoenvironment that was not conducive to habitation by invertebrate organisms. This is consistent with other evidence that Lower Jurassic sauropodomorph tracks are often associated with semi-arid paleoenvironments.

关键词： Sauropodomorph tracks Ziliujing formation Lower Jurassic Guizhou Province

Largest dinosaur tracksite in China (Cretaceous, Zhaojue area, Sichuan Province): On the verge of destruction

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Geoscience Frontiers 2021年第5期12卷 124-137页

作者： Lida Xing Martin G.Lockley hendrik klein W.Scott Persons IV Guangzhao Peng Yong Ye Miaoyan Wang State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing 100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing 100083China Dinosaur Trackers Research Group University of ColoradoDenver 80217USA Saurierwelt Paläontologisches Museum Alte Richt 7D-92318 NeumarktGermany Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA Zigong Dinosaur Museum ZigongSichuan ProvinceChina

The Zhaojue(ZJ)tracksites represent multiple steeply-inclined,track-bearing exposures in the clastic Feitianshan Formation of Sichuan Province that have been sequentially,excavated,mapped,expanded and destroyed,by quarrying,erosion or collapse,for almost three *** quarried area extends for more almost 1.0 km from north to south and-0.5 km from east to *** important track-bearing surfaces have been identified and identified as ZJ-Ⅰ,ZJ-Ⅱ,ZJ-ⅡN and ZJ-Ⅲ,some of which are newly exposed surfaces or expanded surfaces that were previously partially *** represent surfaces lost since they were first reported in papers published in *** ZJ-Ⅱ site represents a hugely expanded continuation of a much smaller track-bearing surface described in *** was mapped using drone technology,and now represents the largest tracksite in China,with a minimum count of 933 recognizable tracks,and the longest recorded sauropod and ornithopod trackways from China(-80 m and-52 m respectively),which also reveal a pronounced change in the direction of trackmaker *** ZJ-Ⅱ surface represents the best Zhaojue paleo-census sample giving a count of 68 individuals from 61 trackways(37 ornithopod,10 theropod,14 sauropod)and isolated tracks(representing 7 individuals).Thus,ornithopods(both large and small trackmakers)represent-54%of the total number of *** smaller ZJ-Ⅲ surface was mapped using traditional methods,and reveals at least 6 ornithopod-and 4 theropod *** the ZJ-Ⅱ and ZJ-Ⅲ surfaces reveal parallel ornithopod trackways suggestive of social or gregarious *** combined data from all four Zhaojue surfaces reveal a total of 1928 tracks,and include a few previously reported pterosaurs and theropod swim *** Zhaojue quarry complex provide a good example of multiple track-bearing sites(surfaces)that require long term study and monitoring to extract as much trackway data as possible before in situ physical evidence is lost.

关键词： Feitianshan Formation Onithopod Teropod Suropod

The Middle-Upper Jurassic dinosaur-pterosaur fauna in Shandong Province,China:Evidence from ichnology

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Journal of Palaeogeography 2023年第3期12卷 419-433页

作者： Li-Da Xing Martin G.Lockley Bo-Lin Tong hendrik klein Chang Liu W.Scott Persons IV State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing100083China Dinosaur Trackers Research Group University of Colorado DenverPO Box 173364DenverCO80217USA Saurierwelt Palaontologisches Museum Alte Richt 7D-92318NeumarktGermany Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharlestonSCUSA

Tracks previously examined between 2002 and 2015 from an incompletely-studied,and now partially lost tracksite site in the Santai Formation of Shandong Province were restudied based on new ex-cavations of additional *** contrast to the previous reports of tridactyl tracks,including Grallator,the present study discovered an additional assemblage of unequivocal pterosaur tracks(Pteraichnus)and a possible didactyl deinonychosaurian *** age of the Santai Formation is controversial but it is probably Late Jurassic,which corresponds to the epoch when pterosaur tracks appear *** interpretations,based on palynology and isotopic data,tentatively suggests an earlier Middle-Late Jurassic age and might suggest that the pterosaurian tracks are as old or somewhat older than any previously reported.

关键词： Santai Formation Theropod Deinonychosaur Pterosaur

Unusual dinosaur trackway preservation as clues to paleo-landscape and behavior from the Lower Cretaceous Luohe Formation,Shaanxi Province,China

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Geoscience Frontiers 2021年第2期12卷 737-745页

作者： Lida Xing Martin G.Lockley Yongzhong Tang hendrik klein W.Scott Persons IV Miaoyan Wang Xingwen Li Hao Wu State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijingChina School of the Earth Sciences and Resources China University of GeosciencesBeijingChina Dinosaur Trackers Research Group University of Colorado DenverPO Box 173364DenverCO 80217USA Shaanxi Institute of Geological Survey Xi'an 710054ShaanxiChina Saurierwelt Paläontologisches Museum Alte Richt 7D–92318 NeumarktGermany Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA

Poorly preserved tracks have limited ichnotaxonomic or biotaxon utility,but may reveal useful information about the paleoenvironment,behavior and track *** mostly parallel to sub parallel trackway segments(T1–T8)were registered on a truncation surface in the Lower Cretaceous Luohe Formation of Shaanxi *** attest to the passage of several bipeds,probably all theropods,in a paleo-contour-parallel,south-north direction in an arid *** of preservation in these trackways is poor,but notably superior in two additional trackways(T9–T10)on a foreset *** T9 indicates a didactyl trackmaker,probably a deinonychosaurian,heading north to *** is the 13th report of deinonychosaurian tracks from the Lower Cretaceous of *** any or all the eight south-north oriented trackway segments represent continuations of other segments in the same trackways,the total number of individual trackmakers heading in this direction may have been as lowas *** the trackway pattern and sedimentological evidence could indicate a physically controlled pathway influencing the direction taken by these trackmakers,the possibility that the trackways also represented small social or gregarious group cannot be ruled out.

关键词： Theropod tracks Ornithischian tracks Deinonychosaurian Dunes sediments

A probable tyrannosaurid track from the Upper Cretaceous of southern China

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Science Bulletin 2019年第16期64卷 1136-1139页

作者： Lida Xing Kecheng Niu Martin G.Lockley hendrik klein Anthony Romilio W.Scott Persons IV Stephen L.Brusatte State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing 100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing 100083China Yingliang Stone Nature History Museum Nan’an 362300China Dinosaur Trackers Research Group University of ColoradoDenver 80217USA Saurierwelt Palaontologisches Museum Neumarkt D-92318Germany School of Biological Sciences the University of QueenslandQueensland 4072Australia Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA School of GeoSciences Grant InstituteUniversity of EdinburghEdinburgh EH9 3FEUK

A large number of Upper Cretaceous(Maastrichtian)dinosaur skeletal fossils have been found in the Nanxiong Basin and adjacent Ganzhou area,including those of theropods,sauropods and ***,known dinosaur ichnoassemblages from the Nanxiong Basin partly indicate a different fauna,with dominating large-and medium-sized ornithopods,theropods,and pterosaurs(1,2)This is one of many known examples where trace and body fossil assemblages from the same formation or geologically discrete region show a different *** and paleobiogeographical analyses therefore should consider data from both,before any conclusions can be *** can be classified according to the relative degree of similarity or congruity(or difference/incongruity)between trace and body fossils into the 5 categories proposed by Lockley[3].

关键词： tyrannosaurid track Upper Cretaceous southern China

Lower Cretaceous sauropod trackways from Lishui City and an overview of dinosaur dominated track assemblages from Zhejiang Province, China

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Journal of Palaeogeography 2018年第4期7卷 326-336页

作者： Li-Da Xing Chun-Yong Chou Martin G.Lockley Anthony Romilio hendrik klein Nan Li W. Scott Persons IV state key laboratory of biogeology and environmental geology china university of geosciencesBeijing 100083China school of the earth sciences and resources china university of geosciencesBeijing 100083China state key laboratory of palaeobiology and stratigraphy nanjing institute of geology and palaeontologychinese academy of sciencesnanjing 210008China dinosaur tracks museum university of colorado denverpo box 173364denverco 80217usa school of biological sciences the university of queenslandbrisbaneqld 4072australia saunerwelt palaontologisches museum alte richt 7d-92318 neumarktgermany department of biological sciences university of alberta11455 saskatchewan driveedmontonab t6g 2e9canada

A relatively small number of tetrapod tracksites from Zhejiang Province fills out the fossil record from that region,which is also known to have yielded both saurischian（titanosauriform, coelurosaur） and ornithischian（basal ornithopod, ankylosaurian） body fossils as well as dinosaurian eggs. We use photogrammetry and 3 D imaging to reinterpret the sauropod tracks from Lishui City, which revealed the existence of trackways. The track record from three documented sites in Zhejiang Province includes avian and non-avian theropod, sauropod, ornithopod and pterosaur tracks. Previous work showed that the purported new bird ichnospecies Dongyangornipes sinensis is a synonym of Uhangrichnus chuni, while we here consider Pteraichnus dongyangensis as a nomen dubium. Such ＂provincial ichnotaxonomy＂ may mask similarities in the ichnofauna across large regions. In fact, we show that the ichnofauna is similar to that found in the Early Cretaceous elsewhere in China.

关键词： Sauropod tracks Theropod tracks Bird tracks Pterosaur tracks Lishui Basin Zhejiang Province

New Early Cretaceous Pterosaur-Bird Track Assemblage from Xinjiang, China:Palaeoethology and Palaeoenvironment

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Acta Geologica Sinica(English Edition) 2013年第6期87卷 1477-1485页

作者： HE Qing XING Lida ZHANG Jianping Martin G.LOCKLEY hendrik klein W.Scott PERSONS IV QI Liqi JIA Chengkai School of the Earth Sciences and Resources China University of Geosciences Dinosaur Tracks Research Group University of Colorado at DenverPO Box 173364 Denver CO 8217 USA Saurierwelt Palontologisches Museum Alte Richt 7 D-92318 Neumarkt Germany Department of Biological Sciences University of Alberta 11455 Saskatchewan DriveEdmonton Alberta T6G 2E9 Canada Research Institute of Experiment and Detection Xinjiang Oilfield Company

A pterosaur-bird track assemblage from a sandstone-siltstone-mudstone sequence of the Lower Cretaceous Tugulu Group of Xinjiang comprises the first pterosaur track record from this province and the largest specimen thus far known from *** pterosaur tracks are assigned to the ichnogenus Pteraichnus based on the triangular overall-shape,the four elongate digit traces and the robust manual digit trace Ⅲ.Supposed trackmakers were dsungaripterid pterodactyloids whose skeletal remains are well known from the Tugulu *** bird tracks that occur on the same surface,are those of typical shorebirds,known from different other localities in southeast *** congruence with Koreanaornis dodsoni described from the same stratigraphic level justifies an assignment to this *** is a further evidence of the co-occurrence of pterosaurs and birds in a typical iakeshore environment with possible seasonal alteration of water supply and aerial exposure indicated by wave ripples,mudcracks and repeated cycles of coarse to fine *** and birds frequented the shoreline and may have fed also on the numerous invertebrates such as the Scoyenia tracemaker that left abundant burrows.

关键词： Pteraichnus Koreanaornis Early Cretaceous Tugulu Group Xinjiang

A new Early Cretaceous dinosaur track assemblage and the first definite non-avian theropod swim trackway from China

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Chinese Science Bulletin 2013年第19期58卷 2370-2378页

作者： XING LiDa LOCKLEY Martin G ZHANG JianPing MILNER Andrew R C klein hendrik LI DaQing PERSONS IV W Scott EBI JieFang School of the Earth Sciences and Resources China University of Geosciences Department of Biological Sciences University of Alberta 11455 Saskatchewan DriveEdmontonAlberta T6G 2E9Canada Dinosaur Tracks Museum University of Colorado Denver St. George Dinosaur Discovery Site at Johnson Farm 2180 East Riverside DriveUtah 84790USA Saurierwelt Pal ontologisches Museum Alte Richt 7D-92318 NeumarktGermany Geological Museum of Gansu Zhaojue County Bureau of Culture MultimediaPress & Sport Tourism

The trackway of a swimming theropod (ichnogenus Characichnos) is reported from the Lower Cretaceous Feitianshan Formation of Sichuan, China. These swim tracks help confirm that non-avian theropods were capable of forging moderately deep bodies of water. The trackway occurs on the same surface as a typical walking trackway of a sauropod (ichnogenus Brontopodus). Both occurrences are the first reported from the Cretaceous of Sichuan, and the swim tracks are the first well-preserved example of a Characichnos trackway from China. Additionally, a theropod walking trackway and several ornithopod walking trackways (similar to the ichnogenus Caririchnium) occur in the same horizon. The ornithopod trackways show a parallel orientation, suggesting gregarious behavior of the trackmakers, which may have been iguanodontiforms and/or hadrosauriforms. The co-occurrence of theropod swim tracks and theropod walking tracks suggests a fluctuation of water depth within a distinct time span.

关键词：蜥脚类恐龙早白垩世兽脚类游泳中国轨道鸟类组合和

The new ichnotaxon Eubrontes nobitai ichnosp. nov. and other saurischian tracks from the Lower Cretaceous of Sichuan Province and a review of Chinese Eubrontes-type tracks

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Journal of Palaeogeography 2021年第3期10卷 302-320页

作者： Li-Da Xing Martin G.Lockley hendrik klein Li-Jun Zhang Anthony Romilio W.Scott Persons IV Guang-Zhao Peng Yong Ye Miao-Yan Wang State Key Laboratory of Biogeology and Environmental Geology China University of GeosciencesBeijing 100083China School of the Earth Sciences and Resources China University of GeosciencesBeijing 100083China Dinosaur Trackers Research Group University of ColoradoDenver 80217USA Saurierwelt Paläontologisches Museum Alte Richt 7 D-92318 NeumarktGermany Institute of Resources and Environment Key Laboratory of Biogenic Traces and Sedimentary Minerals of Henan ProvinceCollaborative Innovation Center of Coalbed Methane and Shale Gas for Central Plains Economic RegionHenan Polytechnic UniversityJiaozuo 454003Henan ProvinceChina School of Biological Sciences The University of QueenslandBrisbaneQueensland 4072Australia Mace Brown Museum of Natural History Department of Geology and Environmental GeosciencesCollege of CharlestonCharleston 29401USA Zigong Dinosaur Museum Zigong 643013Sichuan ProvinceChina

The Jiaguan Formation and the underlying Feitianshan Formation(Lower Cretaceous) in Sichuan Province yield multiple saurischian(theropod–sauropod) dominated ichnofaunas. To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common "generic" labels such as Grallator, Eubrontes, cf. Eubrontes or even "EubrontesMegalosauripus" morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks(Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very wellpreserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.

关键词： Ichnofossils Dinosaur footprints Theropod Myths